K. E. Behre, The Role of Man in European Vegetation History. Vegetation History, pp.633-672, 1988.

B. E. Berglund, The Cultural Landscape during 6000 Years in Southern Sweden, Ecological Bulletins, vol.41, 1991.

B. E. Berglund, H. J. Birks, M. Ralska-jasiewiczowa, and H. E. Wright, Palaeoecological Events during the Last 15,000 Years: Regional Synthesis of Palaeoecological Studies of Lakes and Mires in Europe, 1996.

B. E. Berglund, M. J. Gaillard, L. Bj?-orkman, and T. Persson, Long-term changes in floristic diversity in southern Sweden: palynological richness, vegetation dynamics and land-use, Vegetation History and Archaeobotany, vol.12, issue.2, pp.573-583, 2008.
DOI : 10.1111/j.1365-2699.2005.01290.x

H. J. Birks and H. H. Birks, Biological responses to rapid climate change at the Younger Dryas???Holocene transition at Kr??kenes, western Norway, The Holocene, vol.10, issue.1, 2008.
DOI : 10.1130/0091-7613(2002)030<0971:RAWVRT>2.0.CO;2

H. J. Birks, V. A. Felde, A. E. Bjune, J. Grytnes, H. Sepp?-a et al., Does pollen-assemblage richness reflect floristic richness? A review of recent developments and future challenges. Rev, pp.1-25, 2016.
DOI : 10.1016/j.revpalbo.2015.12.011

L. Bj?-orkman, The establishment of Fagus sylvatica at the stand-scale in southern Sweden, 1999.

J. F. Boyle, M. J. Gaillard, J. O. Kaplan, and J. A. Dearing, Modelling prehistoric land use and carbon budgets, The Holocene, vol.22, issue.5, pp.715-722, 2011.
DOI : 10.1111/j.1365-2486.2008.01712.x

L. R. Bradley, T. Giesecke, K. Halsall, and R. H. Bradshaw, L. outside southern Scandinavia, The Holocene, vol.27, issue.4, pp.579-586, 2013.
DOI : 10.2307/1940219
URL : https://hal.archives-ouvertes.fr/hal-01673005

R. H. Bradshaw and M. Lindbladh, REGIONAL SPREAD AND STAND-SCALE ESTABLISHMENT OF FAGUS SYLVATICA AND PICEA ABIES IN SCANDINAVIA, Ecology, vol.86, issue.7, 2005.
DOI : 10.2307/3543289

A. Brostr?-om, A. Nielsen, M. J. Gaillard, K. L. Hjelle, F. Mazier et al., Pollen productivity estimates of key European plant taxa for quantitative reconstruction of past vegetation: a review, Vegetation History and Archaeobotany, vol.102, issue.5, pp.461-478, 2008.
DOI : 10.1002/jqs.3390010105

C. J. Dalgaard, C. W. Hansen, and N. Kaarsen, Climate Shocks and (Very) Long-run Productivity. Discussion Papers, pp.1-33, 2015.
DOI : 10.2139/ssrn.2665151
URL : http://www.econ.ku.dk/dalgaard/Work/WPs/climateshocks.pdf

B. A. Davis, P. M. Collins, and J. O. Kaplan, The age and post-glacial development of the modern European vegetation: a plant functional approach based on pollen data, Vegetation History and Archaeobotany, vol.314, issue.2, pp.303-317, 2015.
DOI : 10.1126/science.1122667

E. C. Ellis, Anthropogenic transformation of the terrestrial biosphere, Philosophical Transactions of the Royal Society A: Mathematical, Physical and Engineering Sciences, vol.142, issue.2, 2011.
DOI : 10.1016/j.biocon.2008.10.006

W. Finsinger, T. Giesecke, S. Brewer, and M. Leydet, Emergence patterns of novelty in European vegetation assemblages over the past 15??000??years, Ecology Letters, vol.314, issue.3, pp.336-346, 2017.
DOI : 10.1126/science.1122667
URL : https://hal.archives-ouvertes.fr/hal-01681566

R. M. Fyfe, J. L. Beaulieu, H. Binney, R. H. Bradshaw, S. Brewer et al., The European Pollen Database: past efforts and current activities. Veg. Hist. Archaeobotany 18, pp.417-424, 2009.

R. M. Fyfe, C. Twiddle, and S. Sugita, The Holocene vegetation cover of Britain and Ireland: overcoming problems of scale and discerning patterns of openness, Quaternary Science Reviews, vol.73, pp.132-148, 2013.
DOI : 10.1016/j.quascirev.2013.05.014

R. M. Fyfe, J. Woodbridge, and N. Roberts, From forest to farmland: pollen-inferred land cover change across Europe using the pseudobiomization approach, Global Change Biology, vol.41, issue.3, 2015.
DOI : 10.1111/jbi.12353

M. J. Gaillard, S. Sugita, and M. J. Bunting, The use of modelling and simulation approach in reconstructing past landscapes from fossil pollen data: a review and results from the POLLANDCAL network, Vegetation History and Archaeobotany, vol.34, issue.5, pp.419-443, 2008.
DOI : 10.1007/978-3-642-59171-6_18
URL : https://hal.archives-ouvertes.fr/halshs-00961515

M. J. Gaillard, T. Dutoit, K. Hjelle, T. Koff, and M. O-'connell, European cultural landscapes e insights into origins and development, Cultural Landscapes of Europe: Fields of Demeter. Aschenbeck Media, pp.35-44, 2009.

M. J. Gaillard, S. Sugita, and F. Mazier, Holocene land-cover reconstructions for studies on land cover-climate feedbacks, Climate of the Past, vol.6, issue.4, pp.483-499, 2010.
DOI : 10.5194/cp-6-483-2010
URL : https://hal.archives-ouvertes.fr/halshs-00961642

T. Giesecke, T. Hickler, T. Kunkel, M. T. Sykes, and R. H. Bradshaw, ORIGINAL ARTICLE: Towards an understanding of the Holocene distribution of Fagus sylvatica L., Journal of Biogeography, vol.35, issue.1, pp.118-131, 2007.
DOI : 10.14214/sf.481

T. Giesecke and K. D. Bennett, The Holocene spread of Picea abies (L.) Karst. in Fennoscandia and adjacent areas, Journal of Biogeography, vol.41, issue.42, p.31, 2004.
DOI : 10.1017/S0033822200019123

T. Giesecke, B. Davis, and S. Brewer, Towards mapping the late Quaternary vegetation change of Europe, Vegetation History and Archaeobotany, vol.13, issue.1, pp.75-86, 2014.
DOI : 10.1002/(SICI)1099-1417(199803/04)13:2<137::AID-JQS352>3.0.CO;2-6
URL : https://hal.archives-ouvertes.fr/hal-01814536

T. Giesecke, B. Ammann, and A. Brande, Palynological richness and evenness: insights from the taxa accumulation curve, Vegetation History and Archaeobotany, vol.365, issue.3, pp.217-228, 2014.
DOI : 10.1098/rstb.2010.0280

T. Giesecke, S. Brewer, W. Finsinger, M. Leydet, and R. H. Bradshaw, Patterns and dynamics of European vegetation change over the last 15, 2017.
URL : https://hal.archives-ouvertes.fr/hal-01681629

, J. Biogeogr, vol.44, 14411456.

E. Grimm, CONISS: a FORTRAN 77 program for stratigraphically constrained cluster analysis by the method of incremental sum of squares, Computers & Geosciences, vol.13, issue.1, pp.13-35, 1987.
DOI : 10.1016/0098-3004(87)90022-7

J. C. Hargreaves, J. D. Annan, R. Ohgaito, A. Paul, and A. Abe-ouchi, Skill and reliability of climate model ensembles at the Last Glacial Maximum and mid- Holocene. Clim, pp.811-823, 2013.

S. P. Harrison, P. J. Bartlein, K. Izumi, G. Li, J. Annan et al., Evaluation of CMIP5 palaeo-simulations to improve climate projections, Nature Climate Change, vol.20, issue.8, pp.735-743, 2015.
DOI : 10.1029/2009EO160004
URL : https://hal.archives-ouvertes.fr/hal-01806138

T. Hickler, K. Vohland, J. Feehan, P. A. Miller, . Smith et al., Projecting the future distribution of European potential natural vegetation zones with a generalized, tree species-based dynamic vegetation model, Global Ecology and Biogeography, vol.87, issue.1, pp.50-63, 2012.
DOI : 10.1007/s10584-007-9342-4
URL : https://hal.archives-ouvertes.fr/hal-01755868

B. Huntley, P. J. Bartlein, and I. C. Prentice, Climatic Control of the Distribution and Abundance of Beech (Fagus L.) in Europe and North America, Journal of Biogeography, vol.16, issue.6, pp.551-560, 1989.
DOI : 10.2307/2845210

G. L. Jacobson and E. C. Grimm, A Numerical Analysis of Holocene Forest and Prairie Vegetation in Central Minnesota, Ecology, vol.67, issue.4, pp.958-966, 1986.
DOI : 10.2307/1939818

A. M. J?-onsson, F. Lagergren, and . Smith, Forest management facing climate change - an ecosystem model analysis of adaptation strategies, Mitigation and Adaptation Strategies for Global Change, vol.50, issue.1???2, pp.201-220, 2015.
DOI : 10.1023/A:1010663429684

J. O. Kaplan, K. M. Krumhardt, and N. Zimmermann, The prehistoric and preindustrial deforestation of Europe, Quaternary Science Reviews, vol.28, issue.27-28, pp.3016-3034, 2009.
DOI : 10.1016/j.quascirev.2009.09.028

K. Goldewijk, K. K. Beusen, A. Van-drecht, G. De-vos, and M. , The HYDE 3.1 spatially explicit database of human-induced global land-use change over the past 12,000 years, Global Ecology and Biogeography, vol.26, issue.1, pp.73-86, 2011.
DOI : 10.1029/1999GB001232

L. Kullman, Immigration of Picea abies into North-Central Sweden. New evidence of regional expansion and tree-limit evolution, Nordic Journal of Botany, vol.2, issue.1, pp.39-54, 2001.
DOI : 10.2307/3038058

N. Kuosmanen, H. Sepp?-a, T. Alenius, R. H. Bradshaw, J. L. Clear et al., Importance of climate, forest fires and human population size in the Holocene boreal forest composition change in northern Europe, Boreas, vol.1, issue.4, pp.688-702, 2016.
DOI : 10.2307/3543289

N. Kuosmanen, H. Sepp?-a, T. Reitalu, T. Alenius, R. W. Bradshaw et al., Long-term forest composition and its drivers in taiga forest in NW Russia, Vegetation History and Archaeobotany, vol.1, issue.201, pp.221-236, 2016.
DOI : 10.2307/3543289

J. Lechterbeck, K. Edinborough, T. Kerig, R. Fyfe, N. Roberts et al., Is Neolithic land use correlated with demography? An evaluation of pollenderived land cover and radiocarbon-inferred demographic change from Central Europe, Holocene, vol.24, 2014.

P. Legendre, Studying beta diversity: ecological variation partitioning by multiple regression and canonical analysis, Journal of Plant Ecology, vol.21, issue.10, pp.3-8, 2008.
DOI : 10.1126/science.293.5539.2413
URL : http://www.plant-ecology.com/CN/article/downloadArticleFile.do?attachType=PDF&id=6408

L. Roy-ladurie and E. , Histoire humaine et compar ee du climat, 2006.

A. E. Magurran, Measuring Biological Diversity, 2004.

L. Marquer, M. J. Gaillard, and S. Sugita, Holocene changes in vegetation composition in northern Europe: why quantitative pollen-based vegetation reconstructions matter, Quaternary Science Reviews, vol.90, 2014.
DOI : 10.1016/j.quascirev.2014.02.013
URL : https://hal.archives-ouvertes.fr/hal-01174936

I. Matthias, M. Sebastian, S. Semmler, and T. Giesecke, Pollen diversity captures landscape structure and diversity, Journal of Ecology, vol.12, issue.4, pp.880-890, 2015.
DOI : 10.1111/j.1366-9516.2006.00230.x
URL : http://onlinelibrary.wiley.com/doi/10.1111/1365-2745.12404/pdf

A. Mauri, B. A. Davis, P. M. Collins, and J. O. Kaplan, The influence of atmospheric circulation on the mid-Holocene climate of Europe: a dataemodel comparison, Clim. Past, vol.10, 2014.

F. Mazier, M. J. Gaillard, P. Kune-s, S. Sugita, A. K. Trondman et al., Testing the effect of site selection and parameter setting on REVEALS-model estimates of plant abundance using the Czech Quaternary Palynological Database, Review of Palaeobotany and Palynology, vol.187, pp.38-49, 2012.
DOI : 10.1016/j.revpalbo.2012.07.017
URL : https://hal.archives-ouvertes.fr/halshs-00959845

U. Mikolajewicz, M. Gr?-oger, E. Maier-reimer, G. Schurgers, M. Vizca?no et al., Long-term effects of anthropogenic CO2 emissions simulated with a complex earth system model, Climate Dynamics, vol.297, issue.11, pp.599-633, 2007.
DOI : 10.1007/978-3-642-85016-5_23

P. A. Miller, T. Giesecke, T. Hickler, R. H. Bradshaw, . Smith et al., Exploring climatic and biotic controls on Holocene vegetation change in Fennoscandia, Journal of Ecology, vol.6, issue.1, pp.247-259, 2008.
DOI : 10.1029/2004GB002395
URL : http://onlinelibrary.wiley.com/doi/10.1111/j.1365-2745.2007.01342.x/pdf

F. J. Mitchell, How open were European primeval forests? Hypothesis testing using palaeoecological data, Journal of Ecology, vol.22, issue.31, pp.168-177, 2005.
DOI : 10.1016/S0277-3791(96)00037-6

A. B. Nielsen, T. Giesecke, and M. Theuerkauf, Quantitative reconstructions of changes in regional openness in north-central Europe reveal new insights into old questions, Quaternary Science Reviews, vol.47, pp.131-149, 2012.
DOI : 10.1016/j.quascirev.2012.05.011

B. V. Odgaard, Reconstructing past biodiversity, Encyclopedia of Quaternary Science, pp.2508-2514, 2007.

B. Pirzamanbein, J. Lindstr?-om, and A. Poska, Creating spatially continuous maps of past land cover from point estimates: A new statistical approach applied to pollen data, Ecological Complexity, vol.20, pp.127-141, 2014.
DOI : 10.1016/j.ecocom.2014.09.005
URL : https://hal.archives-ouvertes.fr/hal-01175857

I. C. Prentice and R. W. Parsons, Maximum Likelihood Linear Calibration of Pollen Spectra in Terms of Forest Composition, Biometrics, vol.39, issue.4, 1983.
DOI : 10.2307/2531338

M. Ralska-jasiewiczowa, D. Nalepka, and T. Goslar, Some problems of forest transformation at the transition to the oligocratic/ Homo sapiens phase of the Holocene interglacial in northern lowlands of central Europe, Vegetation History and Archaeobotany, vol.12, issue.4, pp.233-247, 2003.
DOI : 10.1007/s00334-003-0021-8

T. Reitalu, H. Sepp?-a, S. Sugita, M. Kangur, T. Koff et al., Long-term drivers of forest composition in a boreonemoral region: the relative importance of climate and human impact, Journal of Biogeography, vol.130, issue.8, 2013.
DOI : 10.1007/978-0-387-87458-6

T. Reitalu, P. Gerhold, A. Poska, M. Partel, V. Vali et al., Novel insights into post-glacial vegetation change: functional and phylogenetic diversity in pollen records, Journal of Vegetation Science, vol.3, issue.5, pp.911-922, 2015.
DOI : 10.1002/ece3.847

G. Schurgers, U. Mikolajewicz, M. Gr?-oger, E. Maier-reimer, M. Vizca?no et al., Dynamics of the terrestrial biosphere, climate and atmospheric CO₂ concentration during interglacials: a comparison between Eemian and Holocene, Climate of the Past, vol.2, issue.2, pp.205-220, 2006.
DOI : 10.5194/cp-2-205-2006
URL : https://www.clim-past.net/2/205/2006/cp-2-205-2006.pdf

H. Sepp?-a, T. Alenius, R. H. Bradshaw, T. Giesecke, M. Heikkil?-a et al., ) and the rise of the boreal ecosystem in Fennoscandia, Journal of Ecology, vol.8, issue.Suppl, pp.629-640, 2009.
DOI : 10.14214/sf.502

S. Shennan, S. S. Downey, A. Timpson, K. Edinborough, S. Colledge et al., Regional population collapse followed initial agriculture booms in mid-Holocene Europe, Nature Communications, vol.32, issue.1, 2013.
DOI : 10.1111/j.1502-3885.2003.tb01227.x
URL : http://www.nature.com/articles/ncomms3486.pdf

P. Smilauer and J. Lep-s, Multivariate Analysis of Ecological Data Using CANOCO 5, 2014.
DOI : 10.1017/CBO9781139627061

B. Smith, I. C. Prentice, and M. T. Sykes, Representation of vegetation dynamics in the modelling of terrestrial ecosystems: comparing two contrasting approaches within European climate space, Global Ecology & Biogeography, vol.10, issue.6, pp.621-637, 2001.
DOI : 10.1046/j.1466-822X.2001.00256.x

D. Smith and N. J. Whitehouse, How fragmented was the British Holocene wildwood? Perspectives on the " Vera " grazing debate from the fossil beetle record, Quat. Sci. Rev, vol.29, pp.539-553, 2010.

W. Steffen, W. Broadgate, L. Deutsch, O. Gaffney, and C. Ludwig, The trajectory of the Anthropocene: The Great Acceleration, The Anthropocene Review, vol.41, issue.1, pp.81-98, 2015.
DOI : 10.1016/B978-0-444-59425-9.00032-9

S. Sugita, Pollen Representation of Vegetation in Quaternary Sediments: Theory and Method in Patchy Vegetation, The Journal of Ecology, vol.82, issue.4, pp.881-897, 1994.
DOI : 10.2307/2261452

S. Sugita, Theory of quantitative reconstruction of vegetation I: pollen from large sites REVEALS regional vegetation composition, The Holocene, vol.9, issue.2, pp.229-241, 2007.
DOI : 10.1016/S0034-6667(03)00137-4

S. Sugita, Theory of quantitative reconstruction of vegetation II: all you need is LOVE, The Holocene, vol.3, issue.2, pp.243-257, 2007.
DOI : 10.1177/0959683607075837

A. K. Trondman, M. J. Gaillard, S. Sugita, R. Fyfe, J. Kaplan et al., Land cover-climate interactions in NW Europe, 6000 BP and 200 BP e first results of the Swedish LANDCLIM project, pp.7-14, 2012.

A. K. Trondman, M. J. Gaillard, and F. Mazier, Pollen-based quantitative reconstructions of Holocene regional vegetation cover (plant-functional types and land-cover types) in Europe suitable for climate modelling, Global Change Biology, vol.87, issue.2, pp.676-697, 2015.
DOI : 10.1007/s10584-007-9342-4
URL : https://hal.archives-ouvertes.fr/hal-01237630

A. K. Trondman, M. J. Gaillard, S. Sugita, L. Bj?-orkman, A. Greisman et al., Are pollen records from small sites appropriate for REVEALS model-based quantitative reconstructions of past regional vegetation? An empirical test in southern Sweden, Vegetation History and Archaeobotany, vol.21, issue.2, pp.131-151, 2016.
DOI : 10.1111/gcb.12737

T. G. Tutin, V. H. Heywood, N. A. Burgess, D. M. Moore, D. H. Valentine et al., Flora Europaea. Volume 3 Diapensiaceae to Myoporaceae, Taxon, vol.22, issue.2/3, 19641980.
DOI : 10.2307/1218149

M. Vellend, Do commonly used indices of ??-diversity measure species turnover?, Journal of Vegetation Science, vol.115, issue.4, pp.545-552, 2001.
DOI : 10.2307/4089518

F. W. Vera, C. N. Waters, J. Zalasiewicz, and C. Summerhayes, Grazing Ecology and Forest History. CAB International The Anthropocene is functionally and stratigraphically distinct from the Holocene, Science, vol.351, pp.138-147, 2000.

K. J. Willis and H. J. Birks, What Is Natural? The Need for a Long-Term Perspective in Biodiversity Conservation, Science, vol.314, issue.5803, 2006.
DOI : 10.1126/science.1122667

J. Woodbridge, R. M. Fyfe, N. Roberts, S. Downey, K. Edinborough et al., The impact of the Neolithic agricultural transition in Britain: a comparison of pollen-based land-cover and archaeological 14C date-inferred population change, Journal of Archaeological Science, vol.51, pp.216-224, 2014.
DOI : 10.1016/j.jas.2012.10.025